Staff+ Application Security Engineer at Anthropic in Remote-Friendly (Travel-Required) | San Francisco

About Anthropic Anthropic’s mission is to create reliable, interpretable, and steerable AI systems. We want AI to be safe and beneficial for our users and for society as a whole. Our team is a quickly growing group of committed researchers, engineers, policy experts, and business leaders working together to build beneficial AI systems. About the role: Anthropic's Application Security team secures the systems that build, serve, and increasingly are Claude. The attack surface is unlike most AppSec work: multi-agent orchestration, sandboxed code execution, agents holding delegated credentials, untrusted tool output crossing trust boundaries — problems with little prior art and no off-the-shelf playbook. The way the team works is also different. We use Claude as our primary tool across every part of the job: it drives our static analysis, drafts and fixes vulnerabilities as pull requests, performs first-line bug bounty triage, and assists threat modeling for design reviews. The human work is the judgment layer — system-level reasoning, deciding what matters, and building the next thing the model can't do yet. This is a builder's role on a senior team. We hire engineers who ship production systems and clear a hands-on threat-modeling bar — people who can find the vulnerability but would rather build the system that finds them all. Every engineer owns a system end-to-end, and…